Kuhry (1988): Intercostal (and costal?) cells with sinuous walls. Stomata arranged at intervals in longitudinal rows of cells lying parallel to the long axis of the leaf. Stomata paracytic, with clear dome-shaped, sometimes almost triangular, subsidiary cells.
Kuhry P (1988) A palaeobotanical and palynological study of Holocene peat from the El Bosque mire. Located in a volcanic area of the Cordillera central of Colombia. Rev. Palaeobot. Palynol. 55:19-72.