Kuhry (1988): Leaves ovate-lanceolate, acuminate, plicate at base, unicostate, the costa ending near the apex, leaf margin serrulate. Leaf cells with papillae projecting from the posterior end of leaf cell wall, narrowly rectangular. Upper leaf cells mostly five times as long as wide. Basal leaf margin with many rows of enlarged cells extended along leaf margin, sometimes up to the widest part of the leaf. Determination following the key of Robinson (1967).
Kuhry P (1988) A palaeobotanical and palynological study of Holocene peat from the El Bosque mire. Located in a volcanic area of the Cordillera central of Colombia. Rev. Palaeobot. Palynol. 55:19-72.