Kuhry (1988): Leaves ovate-lanceolate, longly acuminate, bicostate, the costa ending near mid-leaf, not bordered. Leaf cells smooth, linear.
Kuhry P (1988) A palaeobotanical and palynological study of Holocene peat from the El Bosque mire. Located in a volcanic area of the Cordillera central of Colombia. Rev. Palaeobot. Palynol. 55:19-72.