van Geel 1978: Fruit-bodies 40-122 µm in diameter, darkly pigmented. Wall of covering membrane composed of radiating files of subquadratic cells, 4-7 µm wide at the margin, and narrower at the centre. With a prominent ostiole up to 25 µm long and 6-8 µm wide surrounded by 6-8 hyphae about 2 µm wide. Basal plate with a radiate structure.
van Geel B (1978) A palaeoecological study of Holocene peat bog sections in Germany and the Netherlands, based on the analysis of pollen, spores and macro- and microscopic remains of fungi, algae cormophytes and anymals. Rev. Palaeobot. Palynol. 25:1-120.