van der Wiel (1982): Zygospores square, 36-45 (40) × 32-41 (36) µm, laterally straight. Small pits, about 1 µm in diameter, 3 µm apart are present all over the hyaline surface of the zygospore wall. One to three angles usually retuse.
van der Wiel AM (1982) A palaecological study of a section from the foot of the Hazendonk (Zuid-Holland), based on the analysis of pollen, spores and macroscopic remains. Rev. Palaeobot. Palynol. 38:35-90.