van Geel 1978: sporangia crenulate, 18-40 × 18-32 micron, about 6 micron high, walls about 0.5 micron thick.
Kuhry 1997: Crenulate sporangia, with a central pore.
van Geel B (1978) A palaeoecological study of Holocene peat bog sections in Germany and the Netherlands, based on the analysis of pollen, spores and macro- and microscopic remains of fungi, algae cormophytes and anymals. Rev. Palaeobot. Palynol. 25:1-120.