Non-Pollen Palynomorph Image Database


...“extra fossils“ in pollen slides



The NPP-ID has been successfully transferred to a new hosting at the University of Göttingen. It has now a new domain:
The database can be used as usual with your usual accounts.


The 9th Workshop on Non-Pollen Palynomorphs that will be held 25-28 September 2022 at Adam Mickiewicz University in Poznań (Poland).

It will start on Sunday with the on-site registration and Icebreaker Party. Then it is planned to have a full day of oral and poster presentations. On Tuesday a field trip will take place, followed by dinner. Wednesday will be a day of the microscope session with invited specialists dealing with various organisms – potential NPPs.

If you are interested and would like to join the workshop, write an e-mail to Monika Karpińska-Kołaczek ( by the end of April 2022.


The European Pollen Database (EPD) is planning an in person meeting in Prague in June (1st-3rd) 2022, aiming to bring together palynologists and researchers using different proxies that may be archived in Neotoma (macrofossils, sedaDNA, charcoal, geochemistry, isotopes).

You are invited to training workshops ranging from quantitative land cover reconstructions over Non-Pollen Palynomorphs to several R-based sessions. Topics of keynotes lectures will spread over the diverse proxies and interaction of the Neotoma community including aDNA and archaeobotany. A poster session will give you the opportunity to showcase your own research.

All those interested please complete this form to register. The accommodation of many attendants will be supported, depending on the number of applicants, career stage and country of origin. Registration is free.


Пыльца и НПП из поверхностных проб из Индии опубликованы Quamar and Stivrins (2021).
Dr. Mohd Firoze Quamar любезно предоставил его фотографии НПП для публикации в NPP-ID. Новые НПП представлены в статье без акронимов, поэтому им присвоен акроним NN (no number).
Открытый доступ к фотографиям НПП с разрешения Palynology (10.09.2021).


К пятилетию NPP-ID мы опубликовали статью о базе данных “NPP-ID: Non-Pollen Palynomorph Image Database as a research and educational platform”. Просим цитировать эту статью, если Вы пользуетесь базой данных.
К настоящему времени исследовательская группа NPP-ID объединяет 212 зарегистрированных ученых. База данных дает информацию о 1,635 типах НПП и включает 2,172 фотографий, из которых около 10% находятся в открытом доступе. Они были загружены членами группы или опубликованы в журналах под Creative Commons licences.
Мы предлагаем Вам помочь росту базы данных и загрузить собственные фотографии или рисунки НПП для открытого доступа. В свою очередь мы предлагаем Вам соавторство в следующей статье по развитию базы данных NPP-ID.


Книга “Applications of Non-Pollen Palynomorphs: from Palaeoenvironmental Reconstructions to Biostratigraphy” под редакцией Marret et al. опубликована в Geological Society of London, Special Publications 511.Печатная версия выйдет в августе-сентябре 2021, но все статьи уже доступны онлайн:
O Keefe et al.: Why a new volume on non-pollen palynomorphs?
Shumilovskikh et al.: An overview of the taxonomic groups of non-pollen palynomorphs
Pound et al.: An overview of techniques applied to the extraction of non-pollen palynomorphs, their known taphonomic issues and recommendations to maximize recovery
O Keefe et al.: Nomenclature: how do we designate NPP taxa?
Nuñez Otaño et al.: Palaeomycology: a modern mycological view of fungal palynomorphs
McCarthy et al.: Non-pollen palynomorphs in freshwater sediments and their palaeolimnological potential and selected applications
Andrews et al.: Testate amoebae as non-pollen palynomorphs in pollen slides: usefulness and application in palaeoenvironmental reconstruction
Mudie et al.: Marine dinocysts, acritarchs and less well-known NPP: tintinnids, ostracod and foraminiferal linings, copepod and worm remains
Gauthier & Jouffroy-Bapicot: Detecting human impacts: non-pollen palynomorphs as proxies for human impact on the environment
van Asperen et al.: Coprophilous fungal spores: non-pollen palynomorphs for the study of past megaherbivores
Denison: Stratigraphic and sedimentological aspects of the worldwide distribution of Apectodinium in Paleocene/Eocene Thermal Maximum deposits
Wellman and Ball: Early land plant phytodebris
Agić and Cohen: Non-pollen palynomorphs in deep time: unravelling the evolution of early eukaryotes


Новые НПП были описаны Halbwachs et al (2021) из янтарей под акронимами BAA, BFA и UKA .
Hans Halbwachs любезно предоставил его фотографии НПП для публикации в NPP-ID.
Свободный доступ к фотографиям НПП с разрешения Palynology (31.01.2021).


Глава посвященная НПП анализу только что вышла в книге Amanda Henry Handbook for the Analysis of Micro-Particles in Archaeological Samples.


Усовершенствованная версия NPP Gallery находится он-лайн. Все имеющиеся фотографии базы данных отображаются на Вышем дисплее. Надеемся это облегчит Вашу работу по определению НПП.


НПП на IPC: M04 "Extra microfossils in pollen slides: from environmental indicators to biotic interactions".

Регистрация до 31 марта 2020 на conference website.

Session M04 "Extra microfossils in pollen slides: from environmental indicators to biotic interactions":

Non-pollen palynomorphs (NPPs) represent a large, heterogeneous group of organism remains that appear as “extra microfossils” in pollen slides. Taxonomically, NPPs belong to a wide variety of fungi, animals, algae, and plants. Although many NPPs still remain unknown, they provide essential ecological information that can be used to interpret environmental and landscape changes in the past. NPPs record former environmental conditions, allowing assessment of trophic stage, moisture conditions, dry phases, pastoral activities, disturbances and erosion in the study area. Furthermore, they represent an essential tool for tracing biotic interactions in the last several thousands to millions of years. Identification of NPPs, improvements in understanding their ecological meaning, critical evaluation of their indicative values, biases and limitations of these proxies, are crucial questions for palaeoecological reconstructions. We welcome presenters exploring NPPs in a broad range of disciplines, including modern ecology, palaeoecology, palaeopalynology, forensic science and aerobiology to share their newly gained knowledge.


Новые статьи: - Stivrins N, Cerina A, Gałka M, Heinsalu A, Lõugas L, Veski S (2019) Large herbivore population and vegetation dynamics 14,600–8300 years ago in central Latvia, northeastern Europe. Review of Palaeobotany and Palynology 266: 42-51.
- Stivrins N, Aakala T, Ilvonen L, Pasanen L, Kuuluvainen T, Vasander H, Gałka M, Disbrey HR, Liepins J, Hölmström L, Seppä H (2019) Integrating fire-scar, charcoal and fungal spore data to study fire events in the boreal forest of northern Europe. The Holocene in press DOI: 10.1177/0959683619854524.
- Perrotti AG, van Asperen E (2019) Dung fungi as a proxy for megaherbivores: opportunities and limitations for archaeological applications. Vegetation History and Archaeobotany 28: 93-104.


База данных по НПП функционирует в форме Проекта. Открытый доступ ограничен. Для работы с полной версией базы данных необходима авторизация.


НПП воркшоп планируется во время Pollen Monitoring Program meeting в Риге, 20-23 августа 2019. Регистрация до 20 марта 2019.


Новые статьи по НПП, включенные в литературную базу данных:

Tunno I, Mesing SA (2017) The value of non-pollen palynomorphs in interpreting paleoecological change in the Great Basin (Nevada, USA). Quaternary Research 87: 529-543.
Pound MJ, O Keefe JMK, Nuñez Otaño NB, Riding JB (2019) Three new Miocene fungal palynomorphs from the Brassington Formation, Derbyshire, UK. Palynology, DOI: 10.1080/01916122.2018.1473300.


*** от Encarni Montoya ***

VIII Воркшоп по непыльцевым палиноморфам состоится в Барселоне совместно с рабочей группой по фитолитам с 15 по 19 июля 2019. Места проведения Pompeu Fabra University (UPF) и Insitute of Earth Sciences Jaume ALmera (CSIC).
Секция по фитолитам (15-17 июля) включает ознакомительный воркшоп по работе с фитолитами в археологических и палеоэкологических контекстах. Проводит его Prof. Marco Madella (UPF).
Секция по НПП (17-19 июля) имеет традиционную структуру с устными и постерными докладами, секцией микроскопирования и заключительными дискуссии.
Детали воркшопа можно найти на webpage.


New NPP papers published in 2018 to algal palynomorphs :

McCarthy FMG, Riddick NL, Volik O, Danesh DC, Krueger AM (2018) Algal palynomorphs as proxies of human impact on freshwater resources in the Great Lake region. Anthropocene 21: 16-31.

Stivrins N, Soininen J, Tõnno I, Freiberg R, Veski S, Kisand V (2018) Towards understanding the abundance of non-pollen palynomorphs: A comparison of fossil algae, algal pigments and sedaDNA from temperate lake sediments. Review of Palaeobotany and Palynology 249: 9-15.

A list of papers using NPP in multi-proxy approach :

Enevold R, Rasmussen P, Løvschal M, Olsen J, Odgaard BV (2018) Circumstantial evidence of non-pollen palynomorph palaeoecology: a 5,500 year NPP record from forest hollow sediments compared to pollen and macrofossil inferred palaeoenvironments. Vegetation History and Archaeobotany, in press.

Jones SE, Burjachs F, Ferrer-García C, Giralt S, Schulte L, Fernández-López de Pablo J (2018) A multi-proxy approach to understanding complex responces of salt-lake catchments to climate variability and human pressure: A Late Quaternary case study from south-eastern Spain. Quaternary Science Reviews 184: 201-223.

Bosi G, Labate D, Rinaldi R, Montecchi MC, Mazzanti M, Torri P, Riso FM, Mercuri AM (2018) A survey of the Late Roman period (3rd-6th century AD): Pollen, NPPs and seeds/fruits for reconstructing environmental and cultural changes after the floods in Northern Italy. Quaternary International, in press.


A new NPP gallery is on-line. Please test it and enjoy!


New papers evaluating using of Sporormiella and other NPP in palaeoreconstructions:

Etienne D, Wilhelm B, Sabastier P, Reyss J-L, Arnaud F (2013) Influence of sample location and livestock numbers on Sporormiella concentrations and accumulation rates in surface sediments of Lake Allos, French Alps. Journal of Paleolimnology 49: 117-127.

Etienne D, Jourffroy-Bapicot I (2014) Optimal counting limit for fungal spore abundance estimation using Sporormiella as a case study. Vegetation History and Archaeobotany 23: 743-749.

Doyen E, Etienne D (2017) Ecological and human land-use indicator value of fungal spore morphotypes and assemblages. Vegetation History and Archaeobotany 26: 357-367.


New NPP papers published in 2016 and 2017 and included in the NPP literature database:

Hawksworth DL, van Geel B, Wiltshire PEJ (2016) The enigma of the Diporotheca palynomorph. Review of Palaeobotany and Palynology 235: 94-98.

Revelles J, van Geel B (2016) Human impact and ecological changes in lakeshore environments. The contribution of non-pollen palynomorphs in Lake Banyoles (NE Iberia). Review of Palaeobotany and Palynology 232, 81-97.

Revelles J, Burjachs F, van Geel B (2016) Pollen and non-pollen palynomorphs from the Early Neolithic settlement of La Draga (Girona, Spain). Review of Palaeobotany and Palynology 225, 1-20.

van Asperen EN, Kirby JR, Hunt CO (2016) The effect of preparation methods on dung fungal spores: Implications for recognition of megafaunal populations. Review of Palaeobotany and Palynology 229: 1-8.

Expósito I, Burjachs F, Allué E (2017) Filling in the gaps: The contribution of non-pollen palynomorphs to knowledge about the local environment of the Sierra de Atapuerca caves during the Pleistocene. Quaternary International 433, 224-242.

Ghosh R, Paruya DK, Acharya K, Ghorai N, Bera S (2017) How reliable are non-pollen palynomorphs in tracing vegetation changes and grazing activities? Study from the Darjeeling Himalaya, India. Palaeogeography, Palaeoclimatology, Palaeoecology 475: 23-40.

Musotto LL, Borromei AM, Bianchinotti MV, Coronato A (2017) Late Quaternary palaeoenvironmental reconstruction of central Tierra del Fuego (Argentina) based on pollen and fungi. Quaternary International 442: 13-25.

Rodríguez-Zorro PA, Lima da Costa M, Behling H (2017) Mid-Holocene vegetation dynamics with an early expansion of Mauritia flexuosa palm trees inferred from the Serra do Tepequém in the savannas of Roraima State in Amazonia, northwestern Brazil. Vegetation History and Archaeobotany 26:455-468.

van Asperen E (2017) Fungal diversity on dung of tropical animals in temperate environments: Implications for reconstructing past megafaunal populations. Fungal Ecology 28: 25-32.


NPP session # 28 "Filling the gaps about non-pollen palynomorph ecological significance" at the EPPC 2018 in Dublin, Ireland:
Since first introduced 40 years ago, the description and analysis of terrestrial organic-walled microfossils other than pollen, moss and fern spores from pollen slides have enabled to further explore past environmental conditions, such as relating human activities and landscape. Many of the so called non-pollen palynomorphs have been identified and their ecological significance specified, thus providing valuable additional information to classical palaeo-ecological studies. However, the ecology of many described types is still unknown. The very nature of NPPs is multi-disciplinary. More than ever is it necessary to combine efforts of experts from a broad scale of domains to develop the NPPs’ vast potential.
The objective of this session is to make a leap forward regarding our understanding of the significance of current or new palynomorphs. By comparing the occurrence of new or poorly known NPP taxa with the ecological conditions reconstructed from other proxies, multi-disciplinary studies are likely to reveal the ecological requirements of these types. During this anniversary session, we would like to emphasise such studies in order to shed new light about the ecological value of NPPs. We believe this will serve and extend the interpretation of future studies. To this aim, we welcome proposals that unveil so far unsuspected value to new or known NPPs from any taxonomical group. We encourage proposals covering large time scales within the Quaternary period, but also studies on modern assemblages.

abtract submission


New NPP paper "Non-pollen palynomorphs notes: 2. Holocene record of Megalohypha aqua-dulces, its relation to the fossil form genus Fusiformisporites and association with lignicolous freshwater fungi" by Shumilovskikh et al. is on-line. The link for free download is working until September 02, 2017.


Here is the call for papers for the Association for Environmental Archaeology meeting being held in Edinburgh, UK in December 2017:


The VII Workshop on Non-Pollen Palynomorphs started today. Exciting talks and posters, several microscopic sessions and round-table discussion will take place in Liverpool, UK from 12th to 14th of June 2017. Further details here:


Funding to support early career researchers for participation of the NPP meeting are available:


The VII Workshop on Non-Pollen Palynomorphs will take place in Liverpool, UK from 12th to 14th of June 2017. Deadline for abstract submission is the 14th of May 2017. More information at the workshop webpage:


There is a session on Non-Pollen Palynomorphs (SS16) at the XIV IPC / X IOPC in Salvador (Brazil):
We would be happy to get your contributions!